WX, HW and ZW conceived the idea and designed the study. WX designed the methodology. DX, QW, and HS carried out the searches and organized the occurrence and environmental datasets, performed the analyses, and prepared the figures. WX and HW interpreted the results. WX, HW, and ZW led the writing of the manuscript. WX, HW, TL, and PAB significantly contributed to the manuscript writing and critical review.